PhD Biological & Biomedical Sciences (BBS) at Yale University

Programme overview

The Combined Program in the Biological and Biomedical Sciences (BBS) is the admissions gateway and first-year academic home for students interested in pursuing doctoral degrees in any of the following 12 PhD programs at Yale: Cell Biology; Cellular and Molecular Physiology; Computational Biology and Biomedical Informatics; Experimental Pathology; Genetics; Immunobiology; Interdepartmental Neuroscience Program; Microbiology; Molecular Biophysics and Biochemistry; Molecular, Cellular, and Developmental Biology; Pharmacology; and Program in Translational Biomedicine. Applicants interested in these programs should select BBS on the application and choose the BBS Track which best matches their research interests.

Faculty: Graduate School of Arts and Sciences | Department: Biological & Biomedical Sciences (BBS) | Study mode: full-time | Location: New Haven, CT, United States

Official programme page

Application information

Entry requirements

Standardized tests: GRE is not accepted. English language: TOEFL iBT or IELTS Academic is required of most applicants whose native language is not English. BBS requires a score of at least 600 on the paper version, 250 on the computer-based exam, and 100 on the internet-based exam. Please take the test no later than November and no earlier than 24 months pr Application deadline: December 1 (11:59pm ET).

USD 52,400 a year (2026-27). Graduate School of Arts and Sciences 2026-2027 full-time tuition rate (applies to GSAS programmes). Excludes fees and living costs; PhD students typically receive funding, see the GSAS funding page. Source

PhD applicants at the Graduate School of Arts and Sciences apply for fall 2027 entry; most offers of admission for fall 2027 have a reply deadline of April 15, 2027.

Graduate School of Arts and Sciences deadlines (all 11:59 pm Eastern) for fall 2027: December 1, 2026 (e.g. BBS, Chemistry, Economics, History, Linguistics, Psychology, Public Health, Statistics); December 15, 2026 (e.g. American Studies, Anthropology, Applied Physics, Biomedical Engineering, Computer Science PhD, Electrical & Computer Engineering, Law, Management, Mathematics, Mechanical Engineering, Physics, Political Science, Sociology); January 2, 2027 (e.g. African Studies, Applied and Computational Mathematics, Architecture, Computer Science MS, Earth and Planetary Sciences, East Asian Studies, Nursing, Philosophy). Combined programs follow the earlier deadline or December 15.

The Graduate School of Arts and Sciences application fee for all degree-seeking applicants is US $105, non-refundable, payable by Visa, Mastercard, Discover or American Express; fee waivers can be requested until January 2.

GRE General Test requirements vary by GSAS program (required, optional or not accepted) for the 2026-2027 cycle.

Apply through the official page: https://gsas.yale.edu/admissions/phdmasters-application-process

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